holotype The New York Botanical Garden isotype Herbarium, Royal Botanic Gardens isotype Museum national d'Histoire naturelle isotype Herbario Nacional de Venezuela isotype U. S. National Herbarium isotype Institute of Systematic Botany isotype Herbarium, Missouri Botanical Garden isotype Botanischer Garten und Botanisches Museum Berlin-Dahlem isotype Herbário, Museu Paraense Emílio Goeldi isotype Herbarium, ORSTOM isotype Herbarium, Field Museum of Natural History