Id%Version%Family%Infra family%Hybrid genus%Genus%Infra genus%Hybrid%Species%Species author%Infra species%Rank%Authors%Basionym author%Publishing author%Full name%Full name without family%Full name without authors%Full name without family and authors%Reference%Publication%Collation%Remarks%Hybrid parents%Basionym%Replaced synonym%Distribution%Citation type%Bibliographic reference%Bibliographic type info%Collection date as text%Collection day1%Collection month1%Collection year1%Collection day2%Collection month2%Collection year2%Collection number%Collector team as text%Geographic unit as text%Locality%Latitude degrees%Latitude minutes%Latitude seconds%North or south%Longitude degrees%Longitude minutes%Longitude seconds%East or west%Type remarks%Type name%Type locations%Name status%Original taxon name%Original taxon name author team%Original replaced synonym%Original replaced synonym author team%Original basionym%Original basionym author team%Original parent citation taxon name author team%Original taxon distribution%Original hybrid parentage%Original cited type%Original remarks%Nomenclatural synonym%Other links 34774-1%1.3.1.2%Rubiaceae%%N%Joosia%%N%%%%gen.%H.Karst.%%H.Karst.%Joosia H.Karst.%Joosia H.Karst.%Joosia%Joosia%Fl. Columb. (H. Karst.) 1(1): 9. 1859 [6 Apr 1859] %Fl. Columb. (H. Karst.)%1(1): 9%%%%%%unknown%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%% 44493-1%1.2.2.1.1.2%Rubiaceae%%N%Joosia%sectocalyx%N%%%%subgen.%Steyerm.%%Steyerm.%Joosia subgen. Sectocalyx Steyerm.%Joosia subgen. Sectocalyx Steyerm.%Joosia subgen. Sectocalyx%Joosia subgen. Sectocalyx%Brittonia 27(3): 252. 1975 %Brittonia%27(3): 252%%%%%%unknown%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%% 131689-2%1.3%Rubiaceae%%N%Joosia%%N%aequatoria%%%spec.%Steyerm.%%Steyerm.%Joosia aequatoria Steyerm.%Joosia aequatoria Steyerm.%Joosia aequatoria%Joosia aequatoria%Brittonia 27: 257. 1975 %Brittonia%27: 257%[Gandhi; L. Andersson, Brittonia 49: 38. 1997]%%%%Zamora-Chinchipe (Ecuador, Western South America, Southern America)%tax. nov.%%%%%%%%%%54506%J.A.Steyermark%%%%%%%%%%%%%holotype F%%%%%%%%%%%%%% 77112239-1%1.2%Rubiaceae%%N%Joosia%%N%antioquiana%%%spec.%C.M.Taylor%%C.M.Taylor%Joosia antioquiana C.M.Taylor%Joosia antioquiana C.M.Taylor%Joosia antioquiana%Joosia antioquiana%Novon 20(4): 475 (-476; fig. 2). 2010 [29 Nov 2010] %Novon%20(4): 475 (-476; fig. 2)%%%%%Colombia (Western South America, Southern America)%tax. nov.%%%8 Aug 1985%%%%%%%69%J.G.Ramírez, E.Rentería & O.Mesa%%Antioquia: San Luis, Quebrada La Cristallina, 660-740 m%%%%%%%%%%%holotype JAUM;isotype MO 05017430%%%%%%%%%%%%%% 77175811-1%1.1%Rubiaceae%%N%Joosia%%N%capitata%%%spec.%C.M.Taylor%%C.M.Taylor%Joosia capitata C.M.Taylor%Joosia capitata C.M.Taylor%Joosia capitata%Joosia capitata%Novon 25(2): 222. 2017 [13 Apr 2017] %Novon%25(2): 222%doi:10.3417/2015029%%%%Peru (Western South America, Southern America)%tax. nov.%%%22 Jan 2001%%%%%%%26457%R.Vásquez, R.Rojas, S.Flores & L.Campos%%Cajamarca: prov. San Ignacio, San José de Lourdes, Cerro Las Yeguas, 1900 m%%%%%%%%%%%holotype USM USM-193039;isotype GB GB-0119041;isotype MO MO-4873038%%%%%%%%%%%%%% 77175812-1%1.1%Rubiaceae%%N%Joosia%%N%confusa%%%spec.%C.M.Taylor%%C.M.Taylor%Joosia confusa C.M.Taylor%Joosia confusa C.M.Taylor%Joosia confusa%Joosia confusa%Novon 25(2): 224. 2017 [13 Apr 2017] %Novon%25(2): 224%doi:10.3417/2015029%%%%Peru (Western South America, Southern America);Ecuador (Western South America, Southern America);%tax. nov.%%%22 Mar 1980%%%%%%%4995%M.Rimachi Y.%%Peru: Huánuco: prov. Leoncio Prado, distr. Emileo Baldizan, carretera Tingo María–Pucallpa–La Divisora, c. 1600 m%%%%%%%%%%%holotype USM USM-63673;isotype MO MO-4279717%%%%%%%%%%%%%% 754601-1%1.5.1.2%Rubiaceae%%N%Joosia%%N%dichotoma%%%spec.%(Ruiz & Pav.) H.Karst.%(Ruiz & Pav.)%H.Karst.%Joosia dichotoma (Ruiz & Pav.) H.Karst.%Joosia dichotoma (Ruiz & Pav.) H.Karst.%Joosia dichotoma%Joosia dichotoma%Fl. Columb. (H. Karst.) 1(1): 9. 1859 [6 Apr 1859] %Fl. Columb. (H. Karst.)%1(1): 9%%%%%%unknown%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%(Ruiz & Pav.)%%%%%Cinchona dichotoma%% 131690-2%1.4%Rubiaceae%%N%Joosia%%N%dielsiana%%%spec.%Standl.%%Standl.%Joosia dielsiana Standl.%Joosia dielsiana Standl.%Joosia dielsiana%Joosia dielsiana%Publ. Field Mus. Nat. Hist., Bot. Ser. 11: 219. 1936 %Publ. Field Mus. Nat. Hist., Bot. Ser.%11: 219%[Gandhi; L. Andersson, Brittonia 49: 34. 1997]%%%%Mouth of Rio Santiago, Amazonas (Peru, Western South America, Southern America)%tax. nov.%%%%%%%%%%4478%G.Tessmann%%%%%%%%%%%%%lectotype NY%%%%%%%%%%%%%% 77175813-1%1.1%Rubiaceae%%N%Joosia%%N%frondosa%%%spec.%C.M.Taylor%%C.M.Taylor%Joosia frondosa C.M.Taylor%Joosia frondosa C.M.Taylor%Joosia frondosa%Joosia frondosa%Novon 25(2): 227. 2017 [13 Apr 2017] %Novon%25(2): 227%doi:10.3417/2015029%%%%Colombia (Western South America, Southern America)%tax. nov.%%%17 Oct 1996%%%%%%%3164%R.Sánchez, A.Espitia, C.Moreno & P.Torrijos%%Cauca: mpio. Santa Rosa, aguas arriba desembocadura del Indiyaco en el Caquetá, 380 m%%%%%%%%%%%holotype COAH COAH-032779;isotype COL COL-428662%%%%%%%%%%%%%% 313045-2%1.3%Rubiaceae%%N%Joosia%%N%longisepala%%%spec.%L.Andersson%%L.Andersson%Joosia longisepala L.Andersson%Joosia longisepala L.Andersson%Joosia longisepala%Joosia longisepala%Brittonia 49: 37. 1997 %Brittonia%49: 37%%%%%Pambayacu, Rio Lliquino, 1°29'S, 77°22'W, 420 m, Pastaza (Ecuador, Western South America, Southern America)%tax. nov.%%%%%%%%%%10145%W.A.Palacios%%%%%%%%%%%%%holotype MO;isotype GB;isotype QCNE%%%%%%%%%%%%%% 77175814-1%1.1%Rubiaceae%%N%Joosia%%N%loretensis%%%spec.%(Standl.) C.M.Taylor%(Standl.)%C.M.Taylor%Joosia loretensis (Standl.) C.M.Taylor%Joosia loretensis (Standl.) C.M.Taylor%Joosia loretensis%Joosia loretensis%Novon 25(2): 228. 2017 [13 Apr 2017] %Novon%25(2): 228%doi:10.3417/2015029%%Rubiaceae Rondeletia loretensis Standl.%%%comb. nov.%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%% 131691-2%1.4%Rubiaceae%%N%Joosia%%N%macrocalyx%%%spec.%Standl. ex Steyerm.%%Standl. ex Steyerm.%Joosia macrocalyx Standl. ex Steyerm.%Joosia macrocalyx Standl. ex Steyerm.%Joosia macrocalyx%Joosia macrocalyx%Brittonia 27: 259. 1975 %Brittonia%27: 259%%%%%Napo (Ecuador, Western South America, Southern America)%tax. nov.%%%%%%%%%%2739%F.M.Ownbey%%%%%%%%%%%%%holotype F%%%%%%%%%%%%%% 313041-2%1.3%Rubiaceae%%N%Joosia%%N%multiflora%%%spec.%L.Andersson%%L.Andersson%Joosia multiflora L.Andersson%Joosia multiflora L.Andersson%Joosia multiflora%Joosia multiflora%Brittonia 49: 31. 1997 %Brittonia%49: 31%%%%%Below San Gaban on Rio San Gaban, moist lower montane forest, 500-1000 m, Puno (Peru, Western South America, Southern America)%tax. nov.%%%%%%%%%%1199% A.Herrera, J.Aronson,M.O.Dillon, P.E.Berry%%%%%%%%%%%%%holotype MO;isotype F;isotype NY%%%%%%%%%%%%%% 313046-2%1.3%Rubiaceae%%N%Joosia%%N%obtusa%%%spec.%L.Andersson%%L.Andersson%Joosia obtusa L.Andersson%Joosia obtusa L.Andersson%Joosia obtusa%Joosia obtusa%Brittonia 49: 37. 1997 %Brittonia%49: 37%%%%%Rio Pangui about 10 km S-SE of military camp at Taisha, 2°25'S, 77°28'W, 250-300 m, Morona-Santiago (Ecuador, Western South America, Southern America)%tax. nov.%%%%%%%%%%32109% E.Asanza C.,J.Brandbyge%%%%%%%%%%%%%holotype AAU;isotype GB%%%%%%%%%%%%%% 300965-2%1.3%Rubiaceae%%N%Joosia%%N%oligantha%%%spec.%L.Andersson%%L.Andersson%Joosia oligantha L.Andersson%Joosia oligantha L.Andersson%Joosia oligantha%Joosia oligantha%Fl. Ecuador 50: 76-77 fig. 1994 %Fl. Ecuador%50: 76-77 fig%known only from the type collection%%%%Pastaza (Ecuador, Western South America, Southern America)%tax. nov.%%%%%%%%%%20011%E.Asplund%%%%%%%%%%%%%holotype S%%%%%%%%%%%%%% 131692-2%1.3%Rubiaceae%%N%Joosia%%N%panamensis%%%spec.%Dwyer%%Dwyer%Joosia panamensis Dwyer%Joosia panamensis Dwyer%Joosia panamensis%Joosia panamensis%Ann. Missouri Bot. Gard. 67: 263, fig. 1980 %Ann. Missouri Bot. Gard.%67: 263, fig%[Gandhi; L. Andersson, Brittonia 49: 32. 1997]%%%%Veraguas (Panama, Central America, Southern America)%tax. nov.%%%%%%%%%%25932%T.B.Croat%%%%%%%%%%%%%holotype MO%%%%%%%%%%%%%% 754604-1%1.5%Rubiaceae%%N%Joosia%%N%panamensis%%%spec.%Dwyer%%Dwyer%Joosia panamensis Dwyer%Joosia panamensis Dwyer%Joosia panamensis%Joosia panamensis%Ann. Missouri Bot. Gard. 67(2): 262. 1980 %Ann. Missouri Bot. Gard.%67(2): 262%%%%%%unknown%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%Panama%% 131693-2%1.4%Rubiaceae%%N%Joosia%%N%pulcherrima%%%spec.%Steere%%Steere%Joosia pulcherrima Steere%Joosia pulcherrima Steere%Joosia pulcherrima%Joosia pulcherrima%Flora (Quito) 3: 195. 1943 %Flora (Quito)%3: 195%[Gandhi; L. Andersson, Brittonia 49: 39. 1997]%%%%Ecuador (Western South America, Southern America)%tax. nov.%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%% 77175815-1%1.1%Rubiaceae%%N%Joosia%%N%sericea%%%spec.%(Standl.) C.M.Taylor%(Standl.)%C.M.Taylor%Joosia sericea (Standl.) C.M.Taylor%Joosia sericea (Standl.) C.M.Taylor%Joosia sericea%Joosia sericea%Novon 25(2): 232. 2017 [13 Apr 2017] %Novon%25(2): 232%doi:10.3417/2015029%%Rubiaceae Ladenbergia sericea Standl.%%%comb. nov.%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%%% 131694-2%1.3%Rubiaceae%%N%Joosia%%N%standleyana%%%spec.%Steyerm.%%Steyerm.%Joosia standleyana Steyerm.%Joosia standleyana Steyerm.%Joosia standleyana%Joosia standleyana%Brittonia 27: 257 (-258), figs. 1-7. 1975 %Brittonia%27: 257 (-258), figs. 1-7%[Gandhi; L. Andersson, Brittonia 49: 40. 1997]%%%%Zamora-Chinchipe (Ecuador, Western South America, Southern America)%tax. nov.%%%%%%%%%%54558%J.A.Steyermark%%%%%%%%%%%%%holotype F%%%%%%%%%%%%%% 131695-2%1.3%Rubiaceae%%N%Joosia%%N%ulei%%%spec.%Steyerm.%%Steyerm.%Joosia ulei Steyerm.%Joosia ulei Steyerm.%Joosia ulei%Joosia ulei%Brittonia 27: 255. 1975 %Brittonia%27: 255%[Gandhi; L. Andersson, Brittonia 49: 32. 1997]%%%%Loreto (Peru, Western South America, Southern America)%tax. nov.%%%%%%%%%%6795%E.H.G.Ule%%%%%%%%%%%%%holotype HBG%%%%%%%%%%%%%% 313042-2%1.3.1.2%Rubiaceae%%N%Joosia%%N%umbellifera%%%spec.%H.Karst.%%H.Karst.%Joosia umbellifera H.Karst.%Joosia umbellifera H.Karst.%Joosia umbellifera%Joosia umbellifera%Fl. Columb. (H. Karst.) 1(1): 9, t. 5. 1859 [6 Apr 1859] %Fl. Columb. (H. Karst.)%1(1): 9, t. 5%%%%%Servita, Meta (Colombia, Western South America, Southern America)%tax. nov.%%%%%%%%%%s.n.%G.K.W.H.Karsten, J.J.Triana%%%%%%%%%%%%%lectotype W%%%%%%%%%%%%%% 131696-2%1.3%Rubiaceae%%N%Joosia%%N%umbellifera%H.Karst.%caucana%var.%Steyerm.%%Steyerm.%Joosia umbellifera var. caucana Steyerm.%Joosia umbellifera var. caucana Steyerm.%Joosia umbellifera var. caucana%Joosia umbellifera var. caucana%Brittonia 27: 254. 1975 %Brittonia%27: 254%[Gandhi; L. Andersson, Brittonia 49: 32. 1997]%%%%Cauca (Colombia, Western South America, Southern America)%tax. nov.%%%%%%%%%%1367%E.L.Core%%%%%%%%%%%%%holotype US%%%%%%%%%%%%%% 131697-2%1.3%Rubiaceae%%N%Joosia%%N%umbellifera%%macarenensis%subsp.%Steyerm.%%Steyerm.%Joosia umbellifera subsp. macarenensis Steyerm.%Joosia umbellifera subsp. macarenensis Steyerm.%Joosia umbellifera subsp. macarenensis%Joosia umbellifera subsp. macarenensis%Brittonia 27: 254 (-255)'. 1975 %Brittonia%27: 254 (-255)'%[Gandhi; L. Andersson, Brittonia 49: 32. 1997]%%%%Meta (Colombia, Western South America, Southern America)%tax. nov.%%%%%%%%%%2072% R.Jaramillo Mejia,W.R.Philipson, J.M.Idribo%%%%%%%%%%%%%holotype US%%%%%%%%%%%%%% 131698-2%1.3%Rubiaceae%%N%Joosia%%N%umbellifera%%occidentalis%var.%Steyerm.%%Steyerm.%Joosia umbellifera var. occidentalis Steyerm.%Joosia umbellifera var. occidentalis Steyerm.%Joosia umbellifera var. occidentalis%Joosia umbellifera var. occidentalis%Brittonia 27: 254. 1975 %Brittonia%27: 254%%%%%Colombia (Western South America, Southern America)%tax. nov.%%%%%%%%%%14827%J.Cuatrecasas%%%%%%%%%%%%%holotype US%%%%%%%%%%%%%%